Joel Miller

Role: Protagonist

Background: A hardened survivor and smuggler, Joel lost his daughter Sarah during the outbreak. This trauma left him emotionally closed off and morally ambiguous. Throughout the game, he slowly forms a deep, protective bond with Ellie, viewing her as a second chance at family.

Traits: Tough, resourceful, emotionally scarred, protective.

Ellie Williams

Role: Co-Protagonist

Background: A 14-year-old girl born after the outbreak, Ellie is immune to the Cordyceps infection—a fact that could change the fate of humanity. Despite her age, she's witty, brave, and increasingly capable of handling herself in a dangerous world. Her bond with Joel becomes the emotional core of the story.

Traits: Smart, sarcastic, brave, compassionate, and strong-willed.

Tess

Role: Joel's smuggling partner

Background: Tess is tough, strategic, and fiercely loyal. She initially helps Joel escort Ellie but sacrifices herself early in the journey when she's bitten. Her death is one of the first major losses that drives the story forward.

Traits: Practical, fearless, loyal, no-nonsense.

Tommy Miller

Role: Joel's younger brother

Background: Once a Firefly, Tommy left the group to start a new life. He lives in Jackson, Wyoming, helping build a functioning settlement. He represents a more hopeful, cooperative vision of survival compared to Joel's cynicism.

Traits: Idealistic, loyal, family-oriented, level-headed.

Marlene

Role: Leader of the Fireflies

Background: Marlene is responsible for raising Ellie and leads the Fireflies, a rebel group fighting against the military regime. She believes Ellie's immunity could be humanity's salvation and is willing to sacrifice her for a cure.

Traits: Driven, conflicted, authoritative, pragmatic.